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How fast a given amount of substrate is converted to a product depends on how much enzyme is present By measuring how much product is formed in a given time, the amount of enzyme present can be determined An assay requires that you have some way to determine the concentration of product or substrate at a given time after starting the reaction If the product and substrate have different UV or visible spectra, fluorescence spectra, and so forth, the progress of the reaction can be followed by measuring the change in the spectrum with time If there are no convenient spectral changes, physical separation of substrate and product may be necessary For example, with a radioactive substrate, the appearance of radioactivity in the product can be used to follow product formation

or south of due east or west) The waxing crescent appears in the southwestern or western sky just after sunset and sets 2 to 4 hours after the Sun The Moon at first quarter is in the southern sky at sunset, moves generally westward, and sets around midnight The waxing gibbous Moon is in the southeast at sunset, moves generally westward, and sets in the predawn

Velocity rate, v, activity, d[P]/dt, d[S]/dt is how fast an enzyme converts substrate to product, the amount of substrate consumed, or product formed per unit time Units are micromoles per minute ( mol/min) units

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There are a number of interchangeable words for velocity: the change in substrate or product concentration per time; rate; just plain v (for velocity, often written in italics to convince you it s special); activity; or the calculus equivalent, the first derivative of the product or substrate concentration with respect to time, d[P]/dt or d[S]/dt (the minus means it s going away) Regardless of confusion, velocity (by any of its names) is just how fast you re going Rather than miles per hour, enzyme velocity is measured in molar per minute (M/min) or more usually in micromolar per minute ( M/min)

hours The full Moon is opposite the Sun, rising at sunset and setting at or near sunrise The waning gibbous Moon rises some time after sunset and sets after sunrise the next day The last-quarter Moon rises around midnight and sets around noon The waning crescent waits until the predawn hours to rise and sets in the afternoon Figure 4-5 illustrates the appearance of lunar phases as seen from a midlatitude southern location such as Perth, Australia, or Napier, New Zealand

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Figure 8-1 shows what happens when enzyme is added to a solution of substrate In the absence of enzyme, product appearance is slow, and there is only a small change in product concentration with time (low rate) After enzyme is added, the substrate is converted to product at a much faster rate To measure velocity, you have to actually measure two things: product (or substrate) concentration and time You need a device to measure concentration and a clock Velocity is the slope of a plot of product (or substrate) concentration (or amount) against time Velocity can be expressed in a number of different units The most common is micromolar per minute ( M/min); however, because the velocity depends on the amount of enzyme used in the assay, the velocity is often normalized for the amount of enzyme present by expressing the activity in units of micromoles per minute per milligram of enzyme [ mol/(min mg)] This is called a specific activity You may be wondering (or not) where the volume went after all, product concentration is measured in molar units (M; mol/L) Well, it s really still there, but it

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Figure 4-5 Lunar phases as seen from middle latitudes in the southern hemisphere The Moon s tilt varies somewhat, depending on the season and the time of night

[Product] ( mol/mL)

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time (min)

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